Title: Phylogenetic and population systematics of the Mangabeys (primates: Cercopithecoidea)
Author: Colin P. Groves
On the basis of original craniological studies, it is concluded that recently published claims based on blood proteins that the Mangabeys are diphletic are valid, and it is proposed to divide them into two distinct genera: Cercocebus (with three species: torquatus (including atys), agilis, and galeritus), which belongs to the Cercocebini but has a somewhat isolated position in this tribe, and Lophocebus (with one species, albigena: including aterrimus as a subspecies), which is closer to Papio. The inter-relationships within each genus are discussed, with particular reference to the problem of subspecies vs. allopatric species. A new subspecies is named and described, Lophocebus albigena osmani, and the anomalous position of the poorly-known subspecies L.a. opdenboschi is extensively discussed.
Groves, C.P. (1978). Phylogenetic and population systematics of the mangabeys. Primates 19: 1-34.